Biodiversitas Journal of Biological Diversity
Vol. 27 No. 2 (2026)

Habitat heterogeneity drives fern species diversity in Bilar, Bohol, Philippines using Hill numbers

MA. ANELI A. LANZADERAS (Department of Forestry, College of Forestry and Environmental Science, Bohol Island State University. Bilar Campus, Zamora, Bilar 6317, Bohol, Philippines)
MERI JOYCE SALDOM (Department of Forestry, College of Forestry and Environmental Science, Bohol Island State University. Bilar Campus, Zamora, Bilar 6317, Bohol, Philippines)
ANGELICA P. BALDOS (Department of Forest Science, Faculty of Forestry and Environmental Science, Visayas State University. Visca, Baybay City 6521, Leyte, Philippines)



Article Info

Publish Date
31 May 2026

Abstract

Abstract. Lanzaderas MAA, Saldom MJ, Baldos AP. 2026. Habitat heterogeneity drives fern species diversity in Bilar, Bohol, Philippines using Hill numbers. Biodiversitas 27 (2): d270237. https://doi.org/10.13057/biodiv/d270237. Ferns are ecologically vital components of tropical ecosystems, yet remain underrepresented in biodiversity assessments. This study applied the Hill numbers framework to evaluate the α-diversity of ferns across four habitat types: forest area, hilly terrain, open area, and riparian zone in Bilar, Bohol, Philippines. A total of 24 plots (10 m x 10 m each) were established, with three quadrats per habitat type in each barangay, covering an aggregate area of 2,400 m2. From these plots, we recorded 2,033 individual ferns representing 19 species. Species richness (q = 0) was highest in hilly terrain (13 species), followed by forest area (10), riparian zone (10), and open area (6). At q = 1 and q = 2, the effective numbers of species indicated comparable diversity between hilly terrain, forest area, and riparian zone, while open area showed significantly lower evenness and dominance (p<0.05). Nephrolepis cordifolia and Christella acuminata were the most widespread and abundant taxa, whereas Diplazium esculentum occurred exclusively in riparian zone, underscoring microhabitat specificity. The application of Hill numbers provided a more comprehensive and scale-independent understanding of fern diversity than traditional indices, capturing differences in both species richness and evenness. These findings emphasize that structurally and topographically complex habitats sustain higher effective species numbers. The study advocates prioritizing heterogeneous landscapes in conservation and restoration programs to enhance ecological resilience and safeguard fern diversity in fragmented tropical forests.

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Journal Info

Abbrev

biodiv

Publisher

Subject

Agriculture, Biological Sciences & Forestry Biochemistry, Genetics & Molecular Biology

Description

The Biodiversitas Journal was first published in 2000 by the Department of Biology, FMNS, Universitas Sebelas Maret, Surakarta, Indonesia, then in 2006 it was co-published by the Society for Indonesian Biodiversity and that department; since 2017 it was also hosted by Smujo. From 2003-2012 it was ...