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MUHAMMAD NUR SULTON
Department of Environmental Science, Faculty of Mathematics and Natural Sciences, Universitas Sebelas Maret. Jl. Ir. Sutami 36A, Surakarta 57126, Central Java, Indonesia

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Prediction of potential climate change impacts on the geographic distribution shift of Selaginella kraussiana and S. uncinata in East, South and Southeast Asia AHMAD DWI SETYAWAN; SUTARNO SUTARNO; SUGIYARTO SUGIYARTO; SUNARTO SUNARTO; ILYAS NURSAMSI; MUHAMMAD NUR SULTON; GILANG DWI NUGROHO
Nusantara Bioscience Vol. 18 No. 1 (2026)
Publisher : Smujo International

Show Abstract | Download Original | Original Source | Check in Google Scholar | DOI: 10.13057/nusbiosci/n180105

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Abstract. Setyawan AD, Sutarno, Sugiyarto, Sunarto, Nursamsi I, Sulton MN, Nugroho GD. 2026. Prediction of potential climate change impacts on the geographic distribution shift of Selaginella kraussiana and S. uncinata in East, South and Southeast Asia. Nusantara Bioscience 18 (1): n180105. https://doi.org/10.13057/nusbiosci/n180105. Climate change is increasingly altering the geographic distribution of invasive plant species, yet comparative assessments of closely related invasive taxa remain limited. This study evaluated the potential impacts of climate change on the future distribution of two invasive lycophytes, Selaginella kraussiana and S. uncinata, across East, South, and Southeast Asia. Species distribution models were developed using the Maximum Entropy (MaxEnt) algorithm based on occurrence records compiled from GBIF, field observations, and published literature. The models incorporated 15 environmental variables representing bioclimatic, edaphic, UVB-radiation, and topographic factors. Future habitat suitability was projected for 2030, 2050, and 2080 under four Representative Concentration Pathway (RCP) scenarios (2.6, 4.5, 6.0, and 8.5). Model performance was high for both species, with AUC values of 0.935 for S. kraussiana and 0.966 for S. uncinata, indicating excellent predictive accuracy. Environmental controls differed markedly between the species. Selaginella kraussiana was primarily associated with temperature-related variables, particularly the minimum temperature of the coldest month (bio_6), whereas S. uncinata was more strongly associated with annual precipitation (bio_12) and other moisture-related variables. Future projections indicated substantial habitat expansion for S. kraussiana, with total suitable habitat increasing from 11.47 × 10⁶ km² under current conditions to 16.43 × 10⁶ km² under RCP 8.5 by 2080. Expansion was projected mainly into higher-latitude and higher-elevation subtropical and temperate regions. In contrast, S. uncinata exhibited relatively stable distribution patterns, with total suitable habitat changing only slightly from 2.95 × 10⁶ km² to 3.04 × 10⁶ km² across future climate scenarios. These findings suggest that S. kraussiana may experience a greater climate-driven increase in invasion potential than S. uncinata and demonstrate the value of integrating species distribution modeling with ecological interpretation to support invasion-risk assessment, early detection, and climate-adaptive management of invasive lycophytes in Asia.
Climatic and elevational drivers of Selaginella species richness in Java, Indonesia AHMAD DWI SETYAWAN; SUTARNO SUTARNO; SUGIYARTO SUGIYARTO; SUNARTO SUNARTO; MUHAMMAD NUR SULTON
Nusantara Bioscience Vol. 18 No. 1 (2026)
Publisher : Smujo International

Show Abstract | Download Original | Original Source | Check in Google Scholar | DOI: 10.13057/nusbiosci/n180106

Abstract

Abstract. Setyawan AD, Sutarno, Sugiyarto, Sunarto, Sulton MN. 2026. Climatic and elevational drivers of Selaginella species richness in Java, Indonesia. Nusantara Bioscience 18 (1): n180106. https://doi.org/10.13057/nusbiosci/n180106. Understanding environmental factors that shape species richness patterns, yet the determinants of Selaginella diversity in Java, Indonesia, remain poorly understood. Selaginella, an ancient lycophyte genus widely distributed in tropical regions, is highly sensitive to environmental conditions, particularly moisture availability and temperature. This study examined the relationships between Selaginella species richness and elevational and climatic gradients across Java, Indonesia. A total of 1,962 occurrence records representing 21 identified species and one unidentified taxon were compiled from field surveys, herbarium collections, and verified biodiversity databases. Species richness was estimated using a 0.25° × 0.25° grid system and spatially visualized through Inverse Distance Weighting (IDW) interpolation. The influence of elevation, Annual Mean Temperature, Annual Mean Precipitation, Annual Mean Solar Radiation, and Annual Mean Moisture Index on species richness was evaluated using polynomial regression, Pearson correlation analysis, and Principal Component Analysis (PCA). Species richness showed pronounced spatial heterogeneity, with hotspots concentrated in the humid montane regions of West Java. Richness exhibited nonlinear relationships with all environmental variables and peaked at intermediate elevations. Annual Mean Moisture Index (R² = 0.616) and Annual Mean Precipitation (R² = 0.527) were the variables most strongly associated with species richness, whereas Annual Mean Temperature and solar radiation showed negative relationships. Correlation and PCA results further indicated that moisture availability is closely linked to richness patterns, while elevation influences richness indirectly through its effects on local climatic conditions. These findings demonstrate that humid submontane and montane environments are critical for maintaining Selaginella diversity in Java and highlight the importance of conserving climatically suitable habitats under future environmental change.
Climate-associated shifts in potential habitat of the vulnerable tree Eusideroxylon zwageri in Southeast Asia using an exploratory MaxEnt assessment ARDITA AYU WULANDARI; AULIA TSALATSA AGUSTINA; AZKA DITA AULIA; BIAN RAFI SAFARAZ; ARUM NUR MUKARROMAH; ASFI DZIHNI; ATIKAH KHOIRIYAH AZZAM; AULIA SYAH HAFIFFAH; MUHAMMAD NUR SULTON; AHMAD DWI SETYAWAN
Nusantara Bioscience Vol. 18 No. 1 (2026)
Publisher : Smujo International

Show Abstract | Download Original | Original Source | Check in Google Scholar | DOI: 10.13057/nusbiosci/n180107

Abstract

Abstract. Wulandari AA, Agustina AT, Aulia AD, Safaraz BR, Mukarromah AN, Dzihni A, Azzam AK, Hafiffah AS, Sulton MN, Setyawan AD. 2026. Climate-associated shifts in potential habitat of the vulnerable tree Eusideroxylon zwageri in Southeast Asia using an exploratory MaxEnt assessment. Nusantara Bioscience 18 (1): n180107. https://doi.org/10.13057/nusbiosci/n180107. Eusideroxylon zwageri (Bornean ironwood) is a slow-regenerating, high-value rainforest tree threatened by timber extraction, habitat conversion, and climate change. We examined its current and future potential habitat in Southeast Asia using an archived MaxEnt workflow based on 57 GBIF occurrence records and 18 climatic, topographic, edaphic, vegetation, radiation, and anthropogenic predictors. Baseline climate was represented by WorldClim variables, whereas projections used HadGEM2-ES conditions for 2030, 2050, and 2080 under RCP 2.6, 4.5, 6.0, and 8.5. The baseline model had an AUC of 0.91. Precipitation of the driest quarter contributed 57.40%, followed by human population density (9.85%), minimum temperature of the coldest month (9.43%), and annual temperature range (7.78%). Under baseline conditions, high-suitability habitat occupied 0.22×106 km2 (5.16% of the modeled region), principally in Borneo, Sumatra, Brunei Darussalam, and Peninsular Malaysia. Across future scenarios, total modeled area remained approximately constant (4.25-4.28×106 km2), but the distribution among suitability classes changed; high suitability ranged from 4.94% to 6.82%. Within Borneo, high suitability ranged from 22.35% to 38.06%, and the 2080 suitability centroid shifted 7.81-66.54 km depending on the RCP. These outputs indicate redistribution of modeled environmental suitability, not demonstrated adaptation, occupancy, or population persistence. The maps are therefore appropriate for regional screening but require confirmation through bias-corrected occurrence data, spatially independent validation, multiple climate models, model-tuning and extrapolation diagnostics, dynamic land-cover scenarios, and field assessment before site-level conservation decisions.